Both. What is more, you get to choose. At the bottom of the page there should be a currency option giving you two choices.
An example price listing:
$5.99 when paying with USD
A$9.19 when paying with AUD (note the presence of an A)
Also, if you have a credit card with low-to-no currency conversion fees, I would suggest looking into paying with USD until the price matrix gets updated.
A $5.99 purchase at current visa rates works out at A$8.60.
Another advantage of paying in USD is you avoid GOG's insistence of rounding up every AUD price to end with a 9.